Yellow-browed Warbler
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The yellow-browed warbler (''Phylloscopus inornatus'') is a
leaf warbler Leaf warblers are small insectivorous passerine birds belonging to the genus ''Phylloscopus''. Leaf warblers were formerly included in the Old World warbler family but are now considered to belong to the family Phylloscopidae, introduced in 20 ...
(
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Phylloscopidae) which breeds in the east
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
. This
warbler Various Passeriformes (perching birds) are commonly referred to as warblers. They are not necessarily closely related to one another, but share some characteristics, such as being fairly small, vocal, and insectivorous. Sylvioid warblers Th ...
is strongly migratory and winters mainly in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...
and
South-east Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
, but also in small numbers in western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. Like the rest of Phylloscopidae, it was formerly included in the
Old World warbler Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the ci ...
assemblage. It was formerly considered to comprise three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, but ''P. i. humei'' and ''P. i. mandellii'' are now split as a separate species, Hume's leaf warbler ''P. humei'', leaving ''P. inornatus''
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
. The two
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
differ slightly but consistently in
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
,
bioacoustics Bioacoustics is a cross-disciplinary science that combines biology and acoustics. Usually it refers to the investigation of sound production, dispersion and reception in animals (including humans). This involves neurophysiological and anatomical b ...
, and molecular characters.Svensson, L., Mullarney, K., & Zetterström, D. (2010). ''Collins Bird Guide'', 2nd ed. HarperCollins . Before the species was split, the names yellow-browed willow warbler and inornate warbler were used by a few authors.


Taxonomy

The yellow-browed warbler was first described by the English zoologist
Edward Blyth Edward Blyth (23 December 1810 – 27 December 1873) was an English zoologist who worked for most of his life in India as a curator of zoology at the museum of the Asiatic Society of India in Calcutta. Blyth was born in London in 1810. In 1841 ...
in 1842 and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Regulus inornatus''. The current genus name ''Phylloscopus'' is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''phullon'', "leaf", and ''skopos'', "seeker" (from ''skopeo'', "to watch"). The specific ''inornatus'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "plain".


Description

This is one of the smaller Old World warblers, at 9.5–11 cm long and weighing 4–9 g distinctly smaller than a chiffchaff but slightly larger than
Pallas's leaf warbler Pallas's leaf warbler (''Phylloscopus proregulus'') or Pallas's warbler, is a bird that breeds in mountain forests from southern Siberia east to northern Mongolia and northeast China. It is named for German zoologist Peter Simon Pallas, who fir ...
. Like many other
leaf warbler Leaf warblers are small insectivorous passerine birds belonging to the genus ''Phylloscopus''. Leaf warblers were formerly included in the Old World warbler family but are now considered to belong to the family Phylloscopidae, introduced in 20 ...
s, it has overall greenish upperparts and white underparts. It also has prominent double wing bars formed by yellowish-white tips to the wing covert feathers (a long bar on the greater coverts and a short bar on the median coverts), yellow-margined tertial feathers, and long yellow
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
. Some individuals also have a faint paler green central crown stripe though many do not show this. It is not shy, but its arboreal life style makes it difficult to observe. It is almost constantly in motion. Its song is a high-pitched medley of whistles; the call is piercing, often disyllabic "tseeweest", strikingly loud for the bird's small size. The only real possibility of confusion is with the similar-looking Hume's leaf warbler (''P. humei''), which in the limited area of overlap has duller colours, a faint second wing bar and dark legs and lower mandible. Their songs and calls are clearly distinct, with Hume's having a more chirping "chwee" call. It can easily be distinguished from Pallas's warbler as it does not have the conspicuous yellow central crown stripe and rump patch shown by that species.


Ecology and distribution

Like most warblers, it is insectivorous. The nest is built in dense vegetation often at the base of a tree or old stump; two to four (occasionally more) eggs are laid, hatching after 11–14 days, with the chicks fledging when 12–13 days old. This is an abundant bird of lowland and
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
s and woodlands; particularly in winter it may also be found in more open woods. Its breeding range extends from just west of the
Ural Mountains The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western ...
eastwards to eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
,
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
and
Northeast China Northeast China or Northeastern China () is a geographical region of China, which is often referred to as "Manchuria" or "Inner Manchuria" by surrounding countries and the West. It usually corresponds specifically to the three provinces east of t ...
. Its winter
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
is lowland broadleaf or
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous forest, from
West Bengal West Bengal (, Bengali: ''Poshchim Bongo'', , abbr. WB) is a state in the eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabitants within an area of . West Bengal is the fourt ...
and
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
in northeastern India east through southern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
to
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
, and through
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
south to the
Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area ...
. In summer, it occurs at altitudes of up to 2,440 m, and in winter, up to 1,525 m. The European breeding population west of the Urals has increased westwards in recent decades; in 1950 it was described as 'fairly scarce', but 'locally abundant' with 45,000–46,000 pairs in 1990. Small numbers, most likely from the western end of the breeding range, regularly winter in western Europe. These arrive in
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
in late September and October after a 3,000–3,500 km migration from the Urals, a markedly shorter distance than the 5,500–6,000 km they would need to fly to reach the normal wintering areas in southeastern Asia. Exact numbers in this population are unknown, but typically several hundred are found arriving in Great Britain each autumn; given their unobtrusive behaviour, this is probably only a fraction of the total. In the past widely considered to be
vagrants Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
, these birds are now thought to be undertaking a normal regular migration, able to take advantage of the mild
oceanic climate An oceanic climate, also known as a marine climate, is the humid temperate climate sub-type in Köppen classification ''Cfb'', typical of west coasts in higher middle latitudes of continents, generally featuring cool summers and mild winters ( ...
winters on the western fringes of Europe for wintering. A common species in most of its wide range, the yellow-browed warbler is not considered
threatened Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensat ...
by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
. Hume's leaf warbler overlaps its breeding range with yellow-browed warbler in the western
Sayan Mountains The Sayan Mountains (russian: Саяны ''Sajany''; mn, Соёны нуруу, ''Soyonï nurû''; otk, 𐰚𐰇𐰏𐰢𐰤, Kögmen) are a mountain range in southern Siberia, Russia (Buryatia, Irkutsk Oblast, Krasnoyarsk Krai, Tuva Republic ...
, but the species apparently do not hybridise. Their lineages diverged roughly 2.5
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
.Ernst (1996), Irwin ''et al.'' (2001)


Footnotes


References

* * Ernst, S. (1996): Zweiter Beitrag zur Vogelwelt des Östlichen Altai Second contribution regarding the avifauna of the Sayan Mountains" ''Mitteilungen aus dem Zoologischen Museum Berlin'' 72, ''Suppl. Ann. Ornithol''. 20: 123–180
n German N, or n, is the fourteenth letter in the Latin alphabet, used in the modern English alphabet, the alphabets of other western European languages and others worldwide. Its name in English is ''en'' (pronounced ), plural ''ens''. History ...
* Inskipp, Carol; Inskipp, Tim & Sherub (2000)
The ornithological importance of Thrumshingla National Park, Bhutan.
''
Forktail The forktails are small insectivorous birds in the genus ''Enicurus''. They were formerly in the thrush family, Turdidae, but are more often now treated as part of the Old World flycatcher family, Muscicapidae. Their name derives from their lo ...
'' 14: 147–162. * * *


External links

{{Use dmy dates, date=March 2017
yellow-browed warbler The yellow-browed warbler (''Phylloscopus inornatus'') is a leaf warbler (family Phylloscopidae) which breeds in the east Palearctic. This warbler is strongly migratory and winters mainly in tropical South Asia and South-east Asia, but also ...
Birds of North Asia Birds of Manchuria
yellow-browed warbler The yellow-browed warbler (''Phylloscopus inornatus'') is a leaf warbler (family Phylloscopidae) which breeds in the east Palearctic. This warbler is strongly migratory and winters mainly in tropical South Asia and South-east Asia, but also ...
yellow-browed warbler The yellow-browed warbler (''Phylloscopus inornatus'') is a leaf warbler (family Phylloscopidae) which breeds in the east Palearctic. This warbler is strongly migratory and winters mainly in tropical South Asia and South-east Asia, but also ...